Professor of Biology, Professor of Medical Science


A.B. 1985, Biology, Harvard College
Ph.D. 1991, Organismic and Evolutionary Biology, Harvard University
1990-1993 Junior Fellow, Harvard Society of Fellows
1994-1999 Assistant Professor, University of Massachusetts Amherst
2000-2005 Associate Professor, University of Massachusetts Amherst
2004 Fellow, American Association for the Advancement of Science
2005 Professor, Department of Ecology and Evolutionary Biology, Brown University
2015 Dean’s Award for Excellence in Graduate Mentoring
2016-2019 President, International Society of Vertebrate Morphology
2017-2019 President Elect, Society for Integrative and Comparative Biology

Brown Affiliations

Research Areas

On the Web